ID:LIMK1_HUMAN DESCRIPTION: RecName: Full=LIM domain kinase 1; Short=LIMK-1; EC=2.7.11.1; FUNCTION: Serine/threonine-protein kinase that plays an essential role in the regulation of actin filament dynamics. Acts downstream of several Rho family GTPase signal transduction pathways. Activated by upstream kinases including ROCK1, PAK1 and PAK4, which phosphorylate LIMK1 on a threonine residue located in its activation loop. LIMK1 subsequently phosphorylates and inactivates the actin binding/depolymerizing factors cofilin-1/CFL1, cofilin- 2/CFL2 and destrin/DSTN, thereby preventing the cleavage of filamentous actin (F-actin), and stabilizing the actin cytoskeleton. In this way LIMK1 regulates several actin-dependent biological processes including cell motility, cell cycle progression, and differentiation. Phosphorylates TPPP on serine residues, thereby promoting microtubule disassembly. Stimulates axonal outgrowth and may be involved in brain development. Isoform 3 has a dominant negative effect on actin cytoskeletal changes. CATALYTIC ACTIVITY: ATP + a protein = ADP + a phosphoprotein. SUBUNIT: Interacts (via LIM domain) with the cytoplasmic domain of NRG1 (By similarity). Interacts with NISCH (By similarity). Interacts with RLIM and RNF6 (By similarity). Self-associates to form homodimers. Interacts with HSP90AA1; this interaction promotes LIMK1 dimerization and subsequent transphosphorylation. Interacts with CDN1C. Interacts with SSH1. Interacts with ROCK1. INTERACTION: P29066:Arrb1 (xeno); NbExp=2; IntAct=EBI-444403, EBI-4303019; P29067:Arrb2 (xeno); NbExp=2; IntAct=EBI-444403, EBI-1636616; O95835:LATS1; NbExp=5; IntAct=EBI-444403, EBI-444209; P50281:MMP14; NbExp=4; IntAct=EBI-444403, EBI-992788; Q13153:PAK1; NbExp=5; IntAct=EBI-444403, EBI-1307; Q13177:PAK2; NbExp=2; IntAct=EBI-444403, EBI-1045887; SUBCELLULAR LOCATION: Cytoplasm. Nucleus. Note=Predominantly found in the cytoplasm. TISSUE SPECIFICITY: Highest expression in both adult and fetal nervous system. Detected ubiquitously throughout the different regions of adult brain, with highest levels in the cerebral cortex. Expressed to a lesser extent in heart and skeletal muscle. PTM: Autophosphorylated (By similarity). Phosphorylated on Thr-508 by ROCK1 and PAK1, resulting in activation. Phosphorylated by PAK4 which increases the ability of LIMK1 to phosphorylate cofilin. Phosphorylated at Ser-323 by MAPKAPK2 during activation of VEGFA- induced signaling, which results in activation of LIMK1 and promotion of actin reorganization, cell migration, and tubule formation of endothelial cells. Dephosphorylated and inactivated by SSH1. Phosphorylated by CDC42BP. PTM: Ubiquitinated. 'Lys-48'-linked polyubiquitination by RNF6 leads to proteasomal degradation through the 26S proteasome, modulating LIMK1 levels in the growth cone and its effect on axonal outgrowth. Also polyubiquitinated by RLIM (By similarity). DISEASE: Note=LIMK1 is located in the Williams-Beuren syndrome (WBS) critical region. WBS results from a hemizygous deletion of several genes on chromosome 7q11.23, thought to arise as a consequence of unequal crossing over between highly homologous low-copy repeat sequences flanking the deleted region. SIMILARITY: Belongs to the protein kinase superfamily. TKL Ser/Thr protein kinase family. SIMILARITY: Contains 2 LIM zinc-binding domains. SIMILARITY: Contains 1 PDZ (DHR) domain. SIMILARITY: Contains 1 protein kinase domain.
The RNAfold program from the Vienna RNA Package is used to perform the secondary structure predictions and folding calculations. The estimated folding energy is in kcal/mol. The more negative the energy, the more secondary structure the RNA is likely to have.
ModBase Predicted Comparative 3D Structure on P53667
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Orthologous Genes in Other Species
Orthologies between human, mouse, and rat are computed by taking the best BLASTP hit, and filtering out non-syntenic hits. For more distant species reciprocal-best BLASTP hits are used. Note that the absence of an ortholog in the table below may reflect incomplete annotations in the other species rather than a true absence of the orthologous gene.
AK125156 - Homo sapiens cDNA FLJ43166 fis, clone FCBBF3003373, moderately similar to LIM DOMAIN KINASE 1 (EC 2.7.1.-). AK290162 - Homo sapiens cDNA FLJ75611 complete cds, highly similar to Homo sapiens LIM domain kinase 1 (LIMK1), transcript variant 1, mRNA. D26309 - Homo sapiens LIMK mRNA for LIM kinase, complete cds. JA482219 - Sequence 202 from Patent WO2011072091. JE980511 - Sequence 202 from Patent EP2862929. AB385563 - Synthetic construct DNA, clone: pF1KB8017, Homo sapiens LIMK1 gene for LIM domain kinase 1, complete cds, without stop codon, in Flexi system. BC148340 - Synthetic construct Homo sapiens clone IMAGE:100015326, MGC:182989 LIM domain kinase 1 (LIMK1) mRNA, encodes complete protein. BC152982 - Synthetic construct Homo sapiens clone IMAGE:100016262, MGC:184237 LIM domain kinase 1 (LIMK1) mRNA, encodes complete protein. AF134379 - Homo sapiens LIM domain kinase 1 splice variant (LIMK1) mRNA, complete cds. AK300382 - Homo sapiens cDNA FLJ55661 complete cds, highly similar to LIM domain kinase 1 (EC 2.7.11.1). AK026544 - Homo sapiens cDNA: FLJ22891 fis, clone KAT04587, highly similar to HUMLIMKA Human mRNA for LIMK (LIM kinase). JD128744 - Sequence 109768 from Patent EP1572962. JD104446 - Sequence 85470 from Patent EP1572962. AB209557 - Homo sapiens mRNA for LIM domain kinase 1 isoform 1 variant protein. JD446679 - Sequence 427703 from Patent EP1572962. JD339277 - Sequence 320301 from Patent EP1572962. JD555428 - Sequence 536452 from Patent EP1572962. JD521214 - Sequence 502238 from Patent EP1572962. JD495093 - Sequence 476117 from Patent EP1572962. JD354334 - Sequence 335358 from Patent EP1572962. JD197621 - Sequence 178645 from Patent EP1572962. JD180895 - Sequence 161919 from Patent EP1572962. JD477360 - Sequence 458384 from Patent EP1572962. JD139980 - Sequence 121004 from Patent EP1572962. JD246372 - Sequence 227396 from Patent EP1572962. JD439638 - Sequence 420662 from Patent EP1572962. JD543783 - Sequence 524807 from Patent EP1572962. JD520308 - Sequence 501332 from Patent EP1572962. JD404952 - Sequence 385976 from Patent EP1572962. JD476255 - Sequence 457279 from Patent EP1572962. JD157681 - Sequence 138705 from Patent EP1572962. JD138098 - Sequence 119122 from Patent EP1572962. JD141487 - Sequence 122511 from Patent EP1572962. JD487562 - Sequence 468586 from Patent EP1572962. JD371831 - Sequence 352855 from Patent EP1572962. JD191692 - Sequence 172716 from Patent EP1572962. JD526555 - Sequence 507579 from Patent EP1572962. JD446746 - Sequence 427770 from Patent EP1572962. JD151799 - Sequence 132823 from Patent EP1572962. JD405945 - Sequence 386969 from Patent EP1572962. JD192748 - Sequence 173772 from Patent EP1572962. JD560453 - Sequence 541477 from Patent EP1572962. JD191773 - Sequence 172797 from Patent EP1572962. JD251225 - Sequence 232249 from Patent EP1572962. JD464513 - Sequence 445537 from Patent EP1572962. JD188936 - Sequence 169960 from Patent EP1572962. JD518667 - Sequence 499691 from Patent EP1572962. JD067133 - Sequence 48157 from Patent EP1572962. JD351502 - Sequence 332526 from Patent EP1572962. JD203994 - Sequence 185018 from Patent EP1572962. JD148655 - Sequence 129679 from Patent EP1572962. JD267485 - Sequence 248509 from Patent EP1572962. JD531900 - Sequence 512924 from Patent EP1572962. JD262837 - Sequence 243861 from Patent EP1572962. JD144263 - Sequence 125287 from Patent EP1572962. JD103973 - Sequence 84997 from Patent EP1572962. JD362070 - Sequence 343094 from Patent EP1572962. JD240402 - Sequence 221426 from Patent EP1572962. JD156128 - Sequence 137152 from Patent EP1572962. JD195754 - Sequence 176778 from Patent EP1572962. JD223027 - Sequence 204051 from Patent EP1572962. JD498151 - Sequence 479175 from Patent EP1572962.
Biochemical and Signaling Pathways
BioCarta from NCI Cancer Genome Anatomy Project h_rhoPathway - Rho cell motility signaling pathway h_rac1Pathway - Rac 1 cell motility signaling pathway h_CCR3Pathway - CCR3 signaling in Eosinophils h_malPathway - Role of MAL in Rho-Mediated Activation of SRF
Reactome (by CSHL, EBI, and GO)
Protein P53667 (Reactome details) participates in the following event(s):