ID:MK07_HUMAN DESCRIPTION: RecName: Full=Mitogen-activated protein kinase 7; Short=MAP kinase 7; Short=MAPK 7; EC=2.7.11.24; AltName: Full=Big MAP kinase 1; Short=BMK-1; AltName: Full=Extracellular signal-regulated kinase 5; Short=ERK-5; FUNCTION: Plays a role in various cellular processes such as proliferation, differentiation and cell survival. The upstream activator of MAPK7 is the MAPK kinase MAP2K5. Upon activation, it translocates to the nucleus and phosphorylates various downstream targets including MEF2C. EGF activates MAPK7 through a Ras- independent and MAP2K5-dependent pathway. May have a role in muscle cell differentiation. May be important for endothelial function and maintenance of blood vessel integrity. MAP2K5 and MAPK7 interact specifically with one another and not with MEK1/ERK1 or MEK2/ERK2 pathways. Phosphorylates SGK1 at Ser-78 and this is required for growth factor-induced cell cycle progression. CATALYTIC ACTIVITY: ATP + a protein = ADP + a phosphoprotein. COFACTOR: Magnesium (By similarity). ENZYME REGULATION: Activated by tyrosine and threonine phosphorylation (By similarity). Activated in response to hyperosmolarity, hydrogen peroxide, and epidermal growth factor (EGF). SUBUNIT: Interacts with MAP2K5. Forms oligomers (By similarity). Interacts with MEF2A, MEF2C and MEF2D; the interaction phosphorylates the MEF2s and enhances transcriptional activity of MEF2A, MEF2C but not MEF2D (By similarity). Interacts with SGK1. INTERACTION: Q13163:MAP2K5; NbExp=2; IntAct=EBI-1213983, EBI-307294; P29590:PML; NbExp=6; IntAct=EBI-1213983, EBI-295890; SUBCELLULAR LOCATION: Cytoplasm. Nucleus. Note=Translocates to the nucleus upon activation. TISSUE SPECIFICITY: Expressed in many adult tissues. Abundant in heart, placenta, lung, kidney and skeletal muscle. Not detectable in liver. DOMAIN: The second proline-rich region may interact with actin targeting the kinase to a specific location in the cell. DOMAIN: The TXY motif contains the threonine and tyrosine residues whose phosphorylation activates the MAP kinases. PTM: Dually phosphorylated on Thr-219 and Tyr-221, which activates the enzyme (By similarity). Autophosphorylated in vitro on threonine and tyrosine residues when the C-terminal part of the kinase, which could have a regulatory role, is absent. SIMILARITY: Belongs to the protein kinase superfamily. CMGC Ser/Thr protein kinase family. MAP kinase subfamily. SIMILARITY: Contains 1 protein kinase domain. SEQUENCE CAUTION: Sequence=AAA81381.1; Type=Frameshift; Positions=19, 32; Sequence=BAD92848.1; Type=Erroneous initiation;
The RNAfold program from the Vienna RNA Package is used to perform the secondary structure predictions and folding calculations. The estimated folding energy is in kcal/mol. The more negative the energy, the more secondary structure the RNA is likely to have.
ModBase Predicted Comparative 3D Structure on Q13164
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Orthologous Genes in Other Species
Orthologies between human, mouse, and rat are computed by taking the best BLASTP hit, and filtering out non-syntenic hits. For more distant species reciprocal-best BLASTP hits are used. Note that the absence of an ortholog in the table below may reflect incomplete annotations in the other species rather than a true absence of the orthologous gene.
BC007404 - Homo sapiens mitogen-activated protein kinase 7, mRNA (cDNA clone MGC:2148 IMAGE:3009873), complete cds. BC007992 - Homo sapiens mitogen-activated protein kinase 7, mRNA (cDNA clone MGC:3875 IMAGE:3009873), complete cds. U29725 - Human BMK1 alpha kinase mRNA, complete cds. AK295376 - Homo sapiens cDNA FLJ60404 partial cds, highly similar to Mitogen-activated protein kinase 7 (EC 2.7.11.24). U29726 - Human BMK1 beta kinase mRNA, complete cds. BC030134 - Homo sapiens mitogen-activated protein kinase 7, mRNA (cDNA clone MGC:19581 IMAGE:4111084), complete cds. AB209611 - Homo sapiens mRNA for Mitogen activated protein kinase 7 transcript variant 5 variant protein. BC009963 - Homo sapiens mitogen-activated protein kinase 7, mRNA (cDNA clone MGC:15371 IMAGE:4300124), complete cds. JD195679 - Sequence 176703 from Patent EP1572962. JD141522 - Sequence 122546 from Patent EP1572962. U25278 - Human ERK5 mRNA, complete cds. AY534741 - Homo sapiens mitogen activated protein kinase 7 transcript variant 5 mRNA, complete cds. JD365109 - Sequence 346133 from Patent EP1572962. AB527827 - Synthetic construct DNA, clone: pF1KB5763, Homo sapiens MAPK7 gene for mitogen-activated protein kinase 7, without stop codon, in Flexi system. HQ257997 - Synthetic construct Homo sapiens clone IMAGE:100072306 mitogen-activated protein kinase 7 (MAPK7) gene, encodes complete protein. JD216595 - Sequence 197619 from Patent EP1572962. JD190963 - Sequence 171987 from Patent EP1572962. JD373470 - Sequence 354494 from Patent EP1572962. JD334018 - Sequence 315042 from Patent EP1572962. JD045260 - Sequence 26284 from Patent EP1572962. JD157718 - Sequence 138742 from Patent EP1572962. DM110803 - Analysis System for Expression of Endogenous Antisense RNA. DM110848 - Novel Cancer Marker and Use Thereof. JD309672 - Sequence 290696 from Patent EP1572962. JD371082 - Sequence 352106 from Patent EP1572962. JD334437 - Sequence 315461 from Patent EP1572962. JD189541 - Sequence 170565 from Patent EP1572962. JD245649 - Sequence 226673 from Patent EP1572962.
Biochemical and Signaling Pathways
BioCarta from NCI Cancer Genome Anatomy Project h_erk5Pathway - Role of Erk5 in Neuronal Survival h_hdacPathway - Control of skeletal myogenesis by HDAC & calcium/calmodulin-dependent kinase (CaMK) h_mapkPathway - MAPKinase Signaling Pathway
Reactome (by CSHL, EBI, and GO)
Protein Q13164 (Reactome details) participates in the following event(s):