ID:SIA4C_HUMAN DESCRIPTION: RecName: Full=CMP-N-acetylneuraminate-beta-galactosamide-alpha-2,3-sialyltransferase 4; Short=Alpha 2,3-ST 4; Short=Beta-galactoside alpha-2,3-sialyltransferase 4; EC=2.4.99.-; AltName: Full=Alpha 2,3-sialyltransferase IV; AltName: Full=Gal-NAc6S; AltName: Full=Gal-beta-1,4-GalNAc-alpha-2,3-sialyltransferase; AltName: Full=SAT-3; AltName: Full=ST-4; AltName: Full=ST3Gal IV; Short=ST3GalIV; AltName: Full=ST3GalA.2; AltName: Full=STZ; AltName: Full=Sialyltransferase 4C; Short=SIAT4-C; FUNCTION: It may catalyze the formation of the NeuAc-alpha-2,3- Gal-beta-1,3-GalNAc- or NeuAc-alpha-2,3-Gal-beta-1,3-GlcNAc- sequences found in terminal carbohydrate groups of glycoproteins and glycolipids. It may be involved in the biosynthesis of the sialyl Lewis X determinant. Also acts on the corresponding 1,3- galactosyl derivative. CATALYTIC ACTIVITY: CMP-N-acetylneuraminate + beta-D-galactosyl- 1,4-N-acetyl-D-glucosaminyl-glycoprotein = CMP + alpha-N- acetylneuraminyl-2,3-beta-D-galactosyl-1,4-N-acetyl-D- glucosaminyl-glycoprotein. PATHWAY: Protein modification; protein glycosylation. SUBCELLULAR LOCATION: Golgi apparatus, Golgi stack membrane; Single-pass type II membrane protein. Secreted. Note=Membrane- bound form in trans cisternae of Golgi. Secreted into the body fluid. TISSUE SPECIFICITY: Highly expressed in adult placenta, ovary and testes. PTM: The soluble form derives from the membrane form by proteolytic processing. SIMILARITY: Belongs to the glycosyltransferase 29 family. WEB RESOURCE: Name=GGDB; Note=GlycoGene database; URL="http://riodb.ibase.aist.go.jp/rcmg/ggdb/Homolog?cat=symbol&symbol=ST3GAL4"; WEB RESOURCE: Name=Functional Glycomics Gateway - GTase; Note=ST3Gal IV; URL="http://www.functionalglycomics.org/glycomics/molecule/jsp/glycoEnzyme/viewGlycoEnzyme.jsp?gbpId=gt_hum_625";
The RNAfold program from the Vienna RNA Package is used to perform the secondary structure predictions and folding calculations. The estimated folding energy is in kcal/mol. The more negative the energy, the more secondary structure the RNA is likely to have.
Pfam Domains: PF00777 - Glycosyltransferase family 29 (sialyltransferase)
ModBase Predicted Comparative 3D Structure on Q11206
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Orthologous Genes in Other Species
Orthologies between human, mouse, and rat are computed by taking the best BLASTP hit, and filtering out non-syntenic hits. For more distant species reciprocal-best BLASTP hits are used. Note that the absence of an ortholog in the table below may reflect incomplete annotations in the other species rather than a true absence of the orthologous gene.
E06059 - cDNA encoding alpha 2 to 3 sialyltransferase. LF384537 - JP 2014500723-A/192040: Polycomb-Associated Non-Coding RNAs. X74570 - H.sapiens mRNA for Gal-beta(1-3/1-4)GlcNAc alpha-2.3-sialyltransferase. BC010645 - Homo sapiens ST3 beta-galactoside alpha-2,3-sialyltransferase 4, mRNA (cDNA clone MGC:5017 IMAGE:3449625), complete cds. AK314738 - Homo sapiens cDNA, FLJ95600, Homo sapiens sialyltransferase 4C (beta-galactosidealpha-2,3-sialyltransferase) (SIAT4C), mRNA. AK291577 - Homo sapiens cDNA FLJ75432 complete cds, highly similar to Homo sapiens alpha 2,3-sialyltransferase IV type B+18 mRNA. JD303883 - Sequence 284907 from Patent EP1572962. JD074782 - Sequence 55806 from Patent EP1572962. JD371637 - Sequence 352661 from Patent EP1572962. KJ901738 - Synthetic construct Homo sapiens clone ccsbBroadEn_11132 ST3GAL4 gene, encodes complete protein. EU831879 - Synthetic construct Homo sapiens clone HAIB:100066908; DKFZo004E1022 ST3 beta-galactoside alpha-2,3-sialyltransferase 4 protein (ST3GAL4) gene, encodes complete protein. EU831802 - Synthetic construct Homo sapiens clone HAIB:100066831; DKFZo008E1021 ST3 beta-galactoside alpha-2,3-sialyltransferase 4 protein (ST3GAL4) gene, encodes complete protein. CR456858 - Homo sapiens full open reading frame cDNA clone RZPDo834B1211D for gene SIAT4C, sialyltransferase 4C (beta-galactoside alpha-2,3-sialyltransferase); complete cds, incl. stopcodon. AY040826 - Homo sapiens alpha 2,3-sialyltransferase IV type B+18 mRNA, complete cds; alternatively spliced. AF516604 - Homo sapiens alpha 2,3-sialyltransferase IV type A1-12 mRNA, complete cds; alternatively spliced. AF525084 - Homo sapiens alpha-2,3-sialyltransferase IV type A1+18 mRNA, complete cds; alternatively spliced. L23767 - Human a2,3 sialyltransferse mRNa, complete cds. AF516602 - Homo sapiens alpha 2,3-sialyltransferase IV type A2+18 mRNA, complete cds; alternatively spliced. AF516603 - Homo sapiens alpha 2,3 sialyltransferase IV type A2-12 mRNA, complete cds; alternatively spliced. AK021929 - Homo sapiens cDNA FLJ11867 fis, clone HEMBA1006976, weakly similar to H.sapiens mRNA for Gal-beta(1-3/1-4)GlcNAc alpha-2.3-sialyltransferase. LF375988 - JP 2014500723-A/183491: Polycomb-Associated Non-Coding RNAs. AF035249 - Homo sapiens CMP-N-acetylneuraminate-beta-galactosamide-alpha-2, 3-sialyltransferase mRNA, partial cds. LF375986 - JP 2014500723-A/183489: Polycomb-Associated Non-Coding RNAs. LF375985 - JP 2014500723-A/183488: Polycomb-Associated Non-Coding RNAs. LF375984 - JP 2014500723-A/183487: Polycomb-Associated Non-Coding RNAs. AK128605 - Homo sapiens cDNA FLJ46764 fis, clone TRACH3024428. LF375978 - JP 2014500723-A/183481: Polycomb-Associated Non-Coding RNAs. JD061874 - Sequence 42898 from Patent EP1572962. JD170015 - Sequence 151039 from Patent EP1572962. JD444639 - Sequence 425663 from Patent EP1572962. JD142722 - Sequence 123746 from Patent EP1572962. JD546369 - Sequence 527393 from Patent EP1572962. JD320774 - Sequence 301798 from Patent EP1572962. JD119461 - Sequence 100485 from Patent EP1572962. JD140891 - Sequence 121915 from Patent EP1572962. LF375977 - JP 2014500723-A/183480: Polycomb-Associated Non-Coding RNAs. JD220235 - Sequence 201259 from Patent EP1572962. MA620114 - JP 2018138019-A/192040: Polycomb-Associated Non-Coding RNAs. MA611565 - JP 2018138019-A/183491: Polycomb-Associated Non-Coding RNAs. MA611563 - JP 2018138019-A/183489: Polycomb-Associated Non-Coding RNAs. MA611562 - JP 2018138019-A/183488: Polycomb-Associated Non-Coding RNAs. MA611561 - JP 2018138019-A/183487: Polycomb-Associated Non-Coding RNAs. MA611555 - JP 2018138019-A/183481: Polycomb-Associated Non-Coding RNAs. MA611554 - JP 2018138019-A/183480: Polycomb-Associated Non-Coding RNAs.
Biochemical and Signaling Pathways
BioCyc Knowledge Library PWY-7434 - terminal O-glycans residues modification (via type 2 precursor disaccharide) PWY-7831 - ABH and Lewis epitopes biosynthesis from type 2 precursor disaccharide PWY-7835 - superpathway of glycosphingolipids biosynthesis PWY-7841 - neolacto-series glycosphingolipids biosynthesis
BioCarta from NCI Cancer Genome Anatomy Project h_aMANpathway - Steps in the Glycosylation of Mammalian N-linked Oligosaccarides
Reactome (by CSHL, EBI, and GO)
Protein Q11206 (Reactome details) participates in the following event(s):
R-HSA-1022129 ST3GAL4 transfers Neu5Ac to terminal Gal of N-glycans R-HSA-981497 ST3GAL1-4 can add a sialic acid to the T antigen at the alpha 3 position R-HSA-2046285 The keratan chain can be capped by N-acetylneuraminic acid R-HSA-4084984 ST3GAL1-6 transfer Neu5Ac to terminal Gal (alpha-2,3 link) R-HSA-1912378 Sialylation of Pre-NOTCH R-HSA-975577 N-Glycan antennae elongation R-HSA-977068 Termination of O-glycan biosynthesis R-HSA-2022854 Keratan sulfate biosynthesis R-HSA-4085001 Sialic acid metabolism R-HSA-1912420 Pre-NOTCH Processing in Golgi R-HSA-975576 N-glycan antennae elongation in the medial/trans-Golgi R-HSA-913709 O-linked glycosylation of mucins R-HSA-1638074 Keratan sulfate/keratin metabolism R-HSA-446219 Synthesis of substrates in N-glycan biosythesis R-HSA-1912422 Pre-NOTCH Expression and Processing R-HSA-948021 Transport to the Golgi and subsequent modification R-HSA-5173105 O-linked glycosylation R-HSA-1630316 Glycosaminoglycan metabolism R-HSA-446193 Biosynthesis of the N-glycan precursor (dolichol lipid-linked oligosaccharide, LLO) and transfer to a nascent protein R-HSA-157118 Signaling by NOTCH R-HSA-446203 Asparagine N-linked glycosylation R-HSA-597592 Post-translational protein modification R-HSA-71387 Metabolism of carbohydrates R-HSA-162582 Signal Transduction R-HSA-392499 Metabolism of proteins R-HSA-1430728 Metabolism